Skippers are a group of butterflies placed in the family Hesperiidae within the order Lepidoptera (moths and butterflies). They were previously placed in a separate superfamily, Hesperioidea, but have since been placed in the superfamily Papilionoidea (the butterflies). They are named for their quick, darting flight habits. Most have their antenna tips modified into narrow, hook-like projections. Moreover, skippers mostly lack wing-coupling structure available in most moths. More than 3500 species of skippers are recognized, and they occur worldwide, but with the greatest diversity in the Neotropical regions of Central and South America.
Description and systematics
Traditionally, the Hesperiidae were placed in a monotypic superfamily Hesperioidea, because they are morphologically distinct from other Rhopalocera (butterflies), which mostly belong to the typical butterfly superfamily Papilionoidea. The third and rather small butterfly superfamily was the moth-butterflies (Hedyloidea), which are restricted to the Neotropics, but recent phylogenetic analyses suggest the traditional Papilionoidea are paraphyletic, thus recent classifications treat Hesperiidae and Hedylidae as families within an expanded Papilionoidea to reflect true cladistic relationships.
Grass skipper butterfly (Atalopedes campestris)
Collectively, these three groups of butterflies share many characteristics, especially in the egg, larval, and pupal stages. Nevertheless, skippers have the antennae clubs hooked backward like a crochet hook, while the typical butterflies have club-like tips to their antennae, and moth-butterflies have feathered or pectinate (comb-shaped) antennae similar to moths. Skippers also have generally stockier bodies and larger compound eyes than the other two groups, with stronger wing muscles in the plump thorax, in this resembling many moths more than the other two butterfly lineages do. Unlike, for example, the Arctiinae, though, their wings are usually small in proportion to their bodies....
