Lygodium (climbing fern) is a genus of about 40 species of ferns, native to tropical regions across the world, with a few temperate species in eastern Asia and eastern North America. It is the sole genus in the family Lygodiaceae in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Alternatively, the genus may be placed as the only genus in the subfamily Lygodioideae of a more broadly defined family Schizaeaceae, the family placement used in Plants of the World Online as of November 2019. Per recent molecular evidence, Lygodiaceae is thought to have diverged relatively early from the other members of the Schizaeales due to the relatively high level of synonymous sequence divergence between the families within the Schizaeales.
Description
Lygodium are unusual in that the rachis, or midrib, of the frond is thin, flexible, and long, the frond unrolling with indeterminate growth and the rachis twining around supports, so that each frond forms a distinct vine. The fronds may be from 3–12 m (9.8–39.4 ft) long, depending on the species. They are also easily identifiable by their possession of apical buds that lay dormant until damage to the rachis occurs, allowing them a high degree of endurance.
Range
Lygodium is a wide ranging genus with native populations existing in Asia, Australasia, Africa, and North and South America. The genus is largely pan-tropical, with the center of diversity being Pacific islands, such as Borneo, the Philippine islands, and New Guinea.There do exist several species tolerant of temperate climates such as Lygodium palmatum, which is endemic to the Appalachian region of eastern North America, and Lygodium japonicum, which is native to Japan, but highly invasive in the Southeastern United States. For more on this, refer to the "As invasive species" section below. The lack of extant Lygodium species in Europe is commonly attributed to the Pleistocene glaciation wiping them out. Similar extirpations did not occur in other high...